The compound bud (CB) above the leaf scar
(LS) at each cane node consists of a primary
bud flanked by two secondary buds.
The path to fruitfulness may be
visualized as a series of processes
beginning at bud break the year of bud
formation and ending at bud break the
following year when the buds sprout
and produce flower clusters:
Inflorescence initiation — Formation
of flower clusters begins within
each developing bud when certain tissues
begin to form either an inflorescence
primordia or a tiny tendril.
This
occurs early in the spring, when the
bud is only a few nodes away from the
growing tip.
Plant-mediated necrosis — Some
time after inflorescence initiation in
some buds, the inner section dies above
an apparent abscission layer. This is
clearly a physiological process initiated
by the plant. The plant “decides” that a
bud is not worth the metabolic cost,
and “tells” the bud to die. Secondary
buds on either side of the primary bud
usually survive. This kind of necrosis
seems to peak in the first couple of
months following bloom.
Injury-related necrosis — Injurious
environ- mental factors such as pathogens,
heat, water stress or frost might
damage a bud and cause it to eventually
die.
Bud mites — A strain of the
Erineum mite, Colomerus vitis, grape
bud mites infest only buds. These tiny
worm-shaped mites move from old to
new buds during the first couple of
weeks after bud break, andmultiply all
year. During inspection in the following
winter, we may find hundreds in a
bud, where they can damage or
destroy the inflorescences.
Inflorescence Initiation
The “bud” which occurs at each
node on a cane is actually a compound
bud typically composed of
three simple buds: a primary bud in
the middle and a secondary bud on
each side. For most varieties, under
California conditions, the primary
bud may have up to three flower
clusters, or inflorescences, but more
typically there are two. Thompson
Seedless buds usually have one inflorescence,
sometimes two.
The secondary buds often have no
inflorescence, but sometimes have one
or rarely two; these being smaller than
those in the primary bud. Inside each of
the three buds is a 6- to 12-node compressed
shoot with primordial leaves,
flower clusters and tendrils. Buds that
push at bud break become shoots.1,3,4